E. . the journal of i Ch eck Lis t pase data (a) NOTES ON GEOGRAPHIC DISTRIBUTIO New records of Cryptoblepharus cursor Barbour, 1911 (Squamata, Scincidae, Eugongylinae) from Java, Indonesia Rizal Ananda Sugianto', Djoko Tjahjono Iskandar’, Tati Suryati Syamsudin’ 1 Laboratorium Identifikasi dan Determinasi Il, Sekolah limu dan Teknologi Hayati, Institut Teknologi Bandung, Jalan Ganesa Nomor 10, Bandung 40132, Jawa Barat, Indonesia 2 Laboratorium Analisis Ekosistem Terestrial, Sekolah Ilmu dan Teknologi Hayati, Institut Teknologi Bandung, Jalan Ganesa Nomor 10, Bandung 40132, Jawa Barat, Indonesia Corresponding author: Jati Suryati Syamsudin (tati@itb.ac.id) Abstract. The Lombok Snake-eyed Skink, Cryptoblepharus cursor Barbour, 1911 has been recorded in South Java prior to 1940. However, information on its population on the island are scarce. This study presents new records of C. cursor from Java. Visual Encounter Survey were conducted to observe habitat selections and collect skinks as voucher specimens. South Javan C. cursor are restricted to sandy littoral zones with [pomoea pescaprae (L.) R.Br. — Spinifex littoreus (Burm.f) Merr. whereas Lombok C. cursor are saxicolous, preferring rocky coasts. They also differ from the scansorial C. balinensis Barbour, 1911, a sympatric conge- ner from Java. Key words. Reptile, Snake-eyed Skink, biogeography, vicariance, new record Sugianto RA, Iskandar DT, Syamsudin TS (2024) New records of Cryptoblepharus cursor Barbour, 1911 (Squamata, Scincidae, Eugongylinae) from Java, Indonesia. Check List 20 (4): 833-841. https://doi.org/ 10.15560/20.4.833 INTRODUCTION Cryptoblepharus cursor Barbour, 1911, Lombok Snake-eyed Skink is native to Bali, Lombok, Tengah, and Sangkarang, as well as small islands off Sulawesi, Indonesia, and is associated with coastal habitats (Horner 2007). This skink species was originally thought to be a subspecies to C. boutonii Desjardin, 1831, but it has since been was recognised as a separate species (Greer 1974; Horner 2007). More recently a subspecies of C. cursor, C. c. larsonae Horner, 2007 has been recognised from islands west of Sulawesi (Horner 2007; Koch 2011). However, Brongersma (1940) mentioned snake-eyed skinks have been observed from Muara Bogowonto, an estuary separating Congot and Kadilangu, Daerah Istimewa Yogyakarta. Moreover, our past fieldtrip done in 1976 found skinks in Jawa Barat (= West Java) that are similar to C. cursor. Furthermore, additional reports of skinks identified as C. cursor from Jawa Tengah (= Central Java), Daerah Istimewa Yogyakarta, and Jawa Timur (= East Java) provinces are available in the online citizen-scientist platform iNaturalist (Appendix, Table A1), which support the old literature record. It is possible that these skinks colonised Java by rafting on vegetation debris, as known for other lizards (Rocha et al. 2006). Neverthe- less, reports of C. cursor from outside the published range of the species (Horner 2007; Koch 2011) in Java necessitated investigation. More studies on Cryptoblepharus species, such as C. cursor, are necessary to better understand the OPEN Qaccess ecology and diversity of this widespread genus (Blom et al. 2019). Recent studies by Dodge et al. (2023) and Oliver et al. (2018) have mainly been focused on the conservation of the previously extinct-in-wild C. egeriae Boulenger, 1889, which is endemic to Christmas Island. Horner (2007) and Rocha et al. (2006) men- Aeademicedhon Party Lewood.Jh tion various Cryptoblepharus species in their annotated lists. Cryptoblepharus cursor, both as a recognised Received: 8 April 2024 species or as a subspecies in the past, has only been mentioned few times in the literature: Barbour (1911), Accepted: 18 June 2024 Mertens (1930, 1931), and Horner (2007). Published: 1 July 2024 In this study, we present new records of C. cursor in Jawa Barat, and we confirm the presence of this species in Jawa Tengah and Daerah Istimewa Yogyakarta provinces. These findings provide addi- tional insights into the geographic range of C. cursor and may help in studying the historic pathways of a a ee de ae aves colonisation (Blom et al. 2019) of the genus, which has at least 53 known species (Horner 2007; Uetz rterm if . . A . eae a Re acorn on 40h and Hallermann 2024). We contribute new data to the future study of the cryptic diversity of the genus International — CC BY 4.0) Cryptoblepharus. 833 Sugianto et al. - New records of Cryptoblepharus cursor in Java 834 METHODS Surveys were conducted in the provinces of Daerah Istimewa Yogyakarta and Jawa Tengah from December 2022 to January 2023 and in June 2023; in Jawa Barat, the surveys were conducted in February 2023. Geographic coordinates for each site are provided in Table 1. The westernmost survey site is Batukaras, Jawa Barat, while the easternmost is Karangwuni, Daerah Istimewa Yogyakarta (Figure 1). Distribution maps were created using QGIS Desktop v. 3.34.4 and with the Esri Light Gray Basemap, which is accessible via the HCMGIS v. 241.12 plugin (Quach 2024). We conducted visual encounter surveys to observe and collect skinks and ecological data; we used our bare hands to capture specimens (Inger 1994; Crump and Scott 1994; Fitzgerald 2012; Foster 2012). Collected specimens were fixed in 4% formaldehyde and preserved in 70% ethanol (Foster 2012). Specimen identifications were done based on morphological parameters of C. cursor from primarily Iskandar (unpub- lished) and additionally Horner (2007). The following measurements were recorded for morphometric data: snout—vent length (SVL), tail length (TL), upper arm length (UA), lower arm length (LA), palmar length (HA), forelimb length (FLL), upper leg length (UL), lower leg length (LL), plantar length (FL), and hindlimb length (HLL) following Iskandar (unpublished). Likewise, the following scales were counted to obtain meristic data: scales around thickest part of trunk (BSC), paravertebral scales (vert), ventral scales (vent), subdigital lamellae of fingers and toes 1-5, as well as lamellar formula based on number of subdigital lamellae on each segment of 4" toe (Iskandar unpublished). Prior to their collection, we observed the live skinks for any noteworthy behaviours. All preserved spec- imens from recent surveys have been transferred to Laboratorium Identifikasi dan Determinasi Il, Sekolah Ilmu dan Teknologi Hayati, Institut Teknologi Bandung, Bandung, Jawa Barat, Indonesia under the permit Surat Angkut Tumbuhan dan Satwa - Dalam Negeri S.08/K.22-SKW./SATS-DN/1/2023. RESULTS Cryptoblepharus cursor Barbour, 1911 Figures 2, 3 New records. INDONESIA — DAERAH ISTIMEWA YOGYAKARTA « Kabupaten Kulon Progo, Temon, Congot; 07°54’ 06”S, 110°0212”E; 2-3 m elev.; 11.1.2023; R.A. Sugianto leg.; hand collection; 8 4, 1 9, 16 spec., RX-01 to RX- 08 and RY-01 to RY-08 — JAWA TENGAH « Kabupaten Purworejo, Grabag, Jetis; 07°50’55"S, 109°53’34’E; 1-2 m elev.; 3.VII.2023; R.A. Sugianto obs. — JAWA BARAT « Kabupaten Pangandaran, Cijulang, Batukaras; 07°44’ 09”S, 108°29’54”E; 1-2 m elev.; 18.11.2023; R.A. Sugianto leg.; hand collection; 1 spec., RK-01. Identification. Cryptoblepharus cursor from Java are small, slender skinks with a SVL of 30-45 mm. The tail is longer than the SVL, and some specimens have intact tails reaching over 100 mm in length. Dorsal col- oration is gleaming dark brown or bronze with grey-silver ciliary and dorsal stripes. The venter is pale white or yellowish white (Figures 2, 3A). Few juveniles were observed, each were well developed and appeared as miniature versions of adults without apparent superficial differences. More detailed morphometric and meristic characteristics of C. cursor from Java, as well as a few related species, are presented in Table 2. The new specimens collected in this study are listed in Appendix, Table A2 Table 1. Geographic coordinates of the survey sites. No. Locality Geographic coordinates Jawa Barat, Indonesia 1 Batukaras 07°44'09"S, 108°29'54”E 2 Bojong Salawe 07°42'44"S, 108°30’09"E 3 Batu Hiu 07°41'53”"S, 108°31'04"E 4 Cikembulan 07°41'02"S, 108°36'34”E 5 Pangandaran 07°41'15"S, 108°38’25"E Jawa Tengah, Indonesia 6 Jetis 07°50'55"S, 109°53'34”E Daerah Istimewa Yogyakarta, Indonesia 7 Pasir Mendit 07°53’41"S, 110°01'11”E 8 Kadilangu 07°53'53”"S, 110°01’42”E 9 Congot 07°54'06"S, 110°02'12"E 10 Glagah 07°54’46"S, 110°04’04"E 11 Karangwuni 07°55'14”S, 110°05’03”E Check List 20 (4) - https://doi.org/10.15560/20.4.833 Sugianto et al. - New records of Cryptoblepharus cursor in Java 835 105°0’0,000"E 110°0’0,000"" 115°0'0,000”E 120°0'0,000”E Selat Karimata Kalimantan ny Selat Makassar Belituni ’ i] Sulawesi Sumatra wn LNdonesia 5°0'0,000"S, Jawa a Lombok Legends @ Cryptoblepharus cursor records from Jawa Barat (This study) @ C. cursor records from Jawa Tengah and Daerah Istimewa Yogyakarta (This study) © C. cursor records from Jawa Timur (Iskandar, unpublished) ® C. cursor previous records (Horner 2007; Iskandar unpublished) 0 100 200 km ~~ 105°0’0,000"E 110°0'0,000"E 115°0'0,000"E 120°0'0,000°E 10°0’0,000"S 10°0’0,000"S Samudra Hindia 103°52"30,000°E T1o°00,000°E 110°7°30,000"E Kabupaten Pangandaran, FY Kabupaten Purworejo, Provinsi Jawa Barat, Jawa Tengah, Indonesia Indonesia Kabupaten Kulon Progo, Daerah Istimewa a Yogyakarta, Indonesia Cikembulan e Pangandaran, 6 Batu Hiu PST MAO'S e Pasi . B Salat asir Mendit @, a Kadilangu © Congot ® Glagah @ Karangwuni Samudra Hindia Kabupaten Banyuwangi, Jawa Timur, Indonesia Kabupaten Jembrana, Bali, Indonesia Tengah and ‘ Sangkarang © Selat Bali laut Jawa Laut Flores Figure 1. Locations where C. cursor have been recorded. A. Jawa Barat. B. Jawa 2 ial Tengah and Daerah Istimewa Yogyakarta. beget C. Jawa Timur. D. Previous records of C. cursor (Horner 2007; Iskandar Samudra Hindia PEIN ; 3 Samudra Hindia unpublished). Asien sera Pancur @ Figure 2. Cryptoblepharus cursor specimen RK-01 from Batukaras, Jawa Barat, Indonesia. Check List 20 (4) - https://doi.org/10.15560/20.4.833 Sugianto et al. - New records of Cryptoblepharus cursor in Java 836 Figure 3. Behaviour of Cryptoblepharus cursor at Daerah Istimewa Yogyakarta, Indonesia. A. Basking in the open. B. Hiding under littoral vegetation. C. Hiding under debris. D. Hiding in loose sand. Figure 4. Littoral habitat of Cryptoble- pharus cursor at Karangwuni, Daerah Istimewa Yogyakarta, Indonesia. Geographic range of Cryptoblepharus cursor in Java. Crypoblepharus cursor have been found along the southern shorelines of Jawa Barat, Jawa Tengah, and Daerah Istimewa Yogyakarta, as well as two sites in Java Timur (= East Java) provinces (Figure 1). The population in Jawa Barat was found along a 19 km stretch of shore, and the skinks from Jawa Tengah and Daerah Istimewa Yogyakarta were found along at least 45 km of shore with similar conditions. On the other hand, in Jawa Timur, C. cursor was found at two regions, on the coast of Taman Nasional Alas Purwo at the south-east tip of the island and at a small area at the eastern shore of Kabupaten Banyu- wangi (Iskandar, unpublished; Appendix Table A1). Check List 20 (4) - https://doi.org/10.15560/20.4.833 Sugianto et al. - New records of Cryptoblepharus cursor in Java 837 Table 2. Morphometrics and meristics comparison of select Cryptoblepharus species. SVL = snout-vent length, TL = tail length, UA = upper arm length, LA = lower arm length, PL = palmar length, FLL = forelimb length, UL = upper leg length, LL = lower leg length, FL = foot length, HLL = hindlimb length, BSC = body scale count around midsection, vert = vertebral scales, ven = ventral scales, n" finger or toe = subdigital lamellae count of n" finger or toe. C. cursor C. cursor C. cursor C. balinensis C. virgatus (Jawa Barat) (Daerah Istimewa Yogya- (Jawa Timur) (Jawa Timur) (Tanimbar) karta) Observed data (this study) Comparison data (Iskandar, unpublished) Mean+SD Range n |Mean+SD Range n |Mean+SD _ Range n Mean+SD Range n SVL (mm) 39.440.3 30.0-45.0 47 | 38.3426 325-412 16 | 381423 340-410 7 | 35.0+04 255-397 11 | 35.2403 30.0-38.0 11 TL(mm) 46.7407 34.0*-66.0 43 |41.7+150 67*-618 16 | 516421 480-540 7 | 389408 232-406 6 | 326410 260-450 6 UA/SVL 10.0407 89-125 36 | 110405 98-118 16 | 10.3407 95-115 7 | 13.0409 11.2-147 11 | 9540.3 90-98 11 LA/SVL 8.2+0.7 72-98 44 /101405 98-118 16 | 8241.0 6.7-9.7 7 | 84+0.9 68-95 11 | 75+0.5 67-83 11 PL/SVL 13.641.2 101-163 45 |134+1.0 11.7-15.2 16 | 148408 142-160 7 | 13.7410 12.0-149 11 | 13.4407 12.0-144 11 FLL/SVL 31.6415 28.0-35.0 36 | 345416 313-374 16 | 334422 304-370 7 | 35.0418 318-371 11 | 304412 282-324 10 UL/SVL 12.9407 11.6-14.7 27 | 13.8408 12.4-15.3 16 | 13.0405 12.8-13.7 7 | 141408 12.9-151 11 | 12.3407 11.0-13.2 11 LL/SVL 10.5406 94-117 40 )12.9407 114-144 16 | 10.2407 9.2-11.2 7 9.6 + 1.0 75-111 11: -| 9.6404 8.8-10.1 11 FL/SVL 19.7421 165-229 41 | 21.341.2 19.0-234 16 | 21.441.2 19.7-229 7 | 19.8412 183-214 11 | 18141.0 16.3-19.7 11 HLL/SVL 427+2.0 40.0-48.0 27 | 479422 434-518 16 | 441416 419-460 7 | 436426 387-475 11 | 40.0416 371-425 10 BSC 24.3+1.2 22-26 29 | 22.9+0.9 22-25 = 16 | 24.7+0.5 24-25 7 | 23.7414 20-25 = 11_:-| 21.9403 21-22 11 vert 55.0 + 1.9 51-59 47s | 53.2415 51-56 = 16 «|: 58.1+0.9 57-59 "| Sa 201 49-54 11 - | 479413 46-80 11 ven 64.3+2.3 59-69 24 | 63.9419 60-67 16 | 64.0+1.0 62-65 > | AES 56-65 = 11_:« | 56.5+2.2 54-61 11 1st finger 5.4+0.5 6-7 46 | 67+0.5 6-7 16 | 6.0+0.0 6 7? | 72EOA 7-8 11 | 6.0+0.0 6 11 2nd finger 9.4+0.5 10-11 46 | 10.6+0.5 10-11 16 | 9.7+0.8 9-11 | 12.3205 12-13 11 | 9.2406 8-10 11 3rd finger 12.2+0.8 11-15 46 | 13.5407 12-14 16 | 13.3+1.0 12-15 7 | 15.2+0.8 14-17 1 | 12.4+0.5 12-13 11 Ath finger 13.6411 12-18 45 | 15.3+0.5 15-16 16 | 14.0+0.6 13-15 7 | 176414 15-20 11 | 141406 13-15 11 5th finger 8.4+ 0.7 7-10 46 | 9.2+0.5 8-10 16 | 9140.4 9-10 7 | 11.8+0.4 11-12 11 | 9540.5 9-10 11 Ist toe 7040.3 7-9 42 | 7.7406 7-9 16 | 7.5+0.6 7-8 7)! 8540.8 8-10 11 | 6740.5 6-7 11 2nd toe 11.6 +0.6 12-13 42 | 12.5+40.5 12-13 16 | 11.5+0.6 11-12 7 | 13.6 + 1.0 12-15 11 | 9.8404 9-10 11 3rd toe 16.0+0.9 15-19 42 | 16.9+1.0 15-19 16 | 16.0+0.0 16 7) 187-207 17-19 11 | 14.3+41.0 13-16 11 Ath toe 18.4+0.8 15-21 42 | 20.0+0.9 18-21 16 | 181+0.4 18-19 74) 21. 34-3 20-24 = 11: | 18.4+1.0 17-20 11 oth toe 13.5+0.6 13-16 43 | 13.8411 12-16 16 | 13.9+0.7 13-15 7 | 161411 14-17 11 | 13.6+0.7 12-14 11 Lamellar 5-4-4-5 5-4-3-4 10 | 6-4-4-6 5-4-3-5 16 | 5-3-4-5 5-3-4-5 7 7-4-4-5 7-4-4-5 10 | 5-4-4-4 5-4-4-4 10 formula 6-5-4-6* 6-5-4-6 6-4-5-6 7-5-5-6 6-4-4-5 *Denotes regenerated tail. Habitat preference and behaviour. Cryptob/epharus cursor in Jawa Barat, Jawa Tengah, and Daerah Istimewa Yogyakarta are exclusively found in littoral zones with fine sand and in vegetation dominated by /oomoea pes-caprae L. and/or Spinifex littoreus (Burm.f.) Merr. (Figure 4). The skinks were observed traversing the sand during daytime starting at around 07:00h to 18:00h with peaks occurring at around 10:00h to11:00h and 14:00h to 15:00h. They were often found hiding under littoral vegetation and marine debris (Figure 3B, C). They were rarely observed climbing on debris or lower parts of trees. Furthermore, they were never seen on beaches surrounded by rocky cliffs, whether on the surface or on the outcrops. Sometimes the skink utilize ghost-crab (Ocypodinae) burrows around vegetation or sand-filled pieces of debris as alternative shelters. Lastly, when there are no places to take shelter from perceived threats, these skinks burrow into loose sand (Figure 3D). In contrast, individuals from Jawa Timur C. cursor inhabit rocky coasts and tend to be saxicolous (Bagaskara 2017; Iskandar unpublished), similar to their counterparts from Lombok (Horner 2007; Iskandar unpublished). Javan C. cursor in the wild were not observed feeding on other organisms nor they were observed being consumed on by predators during the survey. DISCUSSION Colonisation of Java by Cryptoblepharus cursor. Cryptoblepharus cursor are now found in the southern coasts of Jawa Barat, Jawa Tengah, Daerah Istimewa Yogyakarta, and two sites at Jawa Timur and their predicted range is likely limited to these shorelines (Figure 1), given their observed habitat preference. The subpopulation in Jawa Barat is separated by approximately 800 km from the westernmost tip of Lombok. Check List 20 (4) - https://doi.org/10.15560/20.4.833 Sugianto et al. - New records of Cryptoblepharus cursor in Java 838 However, the subpopulation throughout Jawa Tengah and Daerah Istimewa Yogyakarta is at least about 620 km distant from Lombok. The two sites inhabited by C. cursor in Jawa Timur, meanwhile, are closer to Lombok, which is about 160 km away. Cryptoblepharus cursor is known to traverse coastal habitats (Horner 2007; Iskandar unpublished). This species and C. keiensis Roux, 1910 are similar in this regard, although the former may venture deeper inland (Iskandar unpublished). Cryptoblepharus cursor might have colonised the islands by rafting, a pathway that allows for oceanic dispersal of terrestrial reptiles (Rocha et al. 2006; Reilly et al. 2019). It is also possible that C. cursor spread to Java through human-mediated introduction events, like some other reptile species in the archipelago (Reilly et al. 2019). Five species belonging to order Squamata, namely the Lycodon capucinus Boie, 1827, Lycodon subcinctus Boie, 1827, Trimeresurus insularis Kramer, 1877, Eutropis multifasciata (Kuhl, 1820), and Gekko gecko (Linnaeus, 1758), have spread throughout Nusa Tenngara, either through natural or anthropogenic means, in relatively recent times (Reilly et al. 2019). Nevertheless, without known nucleotide diversity of Cryptoblepharus of Java to determine the phylogenetic relationships and ages of these species, the true nature of their colonisation cannot be ascertained as of now. Natural history of Javan Cryptoblepharus cursor. Brongersma (1940) mentioned that Felix Kopstein obseved snake-eyed skinks from Daerah Istimewa Yogyakarta and later, our previous surveys in 1976 found snake-eyed skinks similar to C. cursor in Jawa Barat and Jawa Timur Our past fieldtrip made a single obser- vation along approximately 19 km of the southern shoreline of Jawa Barat and our February 2023 fieldtrip resurveyed these areas. The sandy shorelines inhabited by the skinks from Jawa Barat, Jawa Tengah, and Daerah Istimewa Yogyakarta span 20-100 m wide and some of the areas were frequently visited by tourists, fishermen, and aquaculture caretakers, while in other areas, human activities were negligible. The area to the west of Batukaras, Jawa Barat, consists of vertical rocky outcrops, and is not suitable habitat for C. cursor. The sand-dwelling of C. cursor from the three main survey areas were only found in the intertidal zone, especially where some marine debris is present. The habitat of this species is dominated by /pomoea pes-caprae—Spinifex littoreus formations (Figure 4). This skink species has never been observed out- side the intertidal zone, which is similar to the coast-inhabiting C. schlegelianus Mertens, 1928 of Timor (Brongersma 1940), C. balinensis sumbawanus Mertens, 1928 (Mertens 1928, 1930, 1931), and perhaps C. richardsi Horner, 2007 of Papua (Shea and Tallowin 2015). Cryptoblepharus cursor from Jawa Timur differs in habitat selection compared to the south Javan subpopulations in that it is saxicolous and readily climbs rocks like their counterparts from Lombok (Horner 2007; Iskandar unpublished). Cryptoblepharus cursor may also climb low vegetation, an individual from Batukaras, Jawa Barat during the 2023 survey was seen climbing the trunk of a tree about 30 cm from the ground. Moreover, C. cursor near Sangsit, Bali were observed living in proximity to human habitations and on tree trunks, which is consistent with Horner (2007). Nevertheless, the collective behaviour and habitat preference of C. cursor differs from C. balinensis, another species from Java. Cryptoblepharus balinensis from around Surabaya and Madura, Jawa Timur, as well as on Bali, has been observed to climb trees, similar to C. keiensis from Maluku (Iskandar unpublished) and C. pannosus Horner, 2007, formerly known as C. carnabyi (Borner and Schittler 1981)..Cryptoblepharus balinensis additionally have been found up to 30 km inland in Jawa Timur and up to 800 m above sea level in Bali (Iskandar, unpublished). Our observations suggest that C. cursor from Jawa Barat, Jawa Tengah, and Daerah Istimewa Yogyakar- ta start their activity around 07:00h and conclude at 18:00h with major peaks at approximately 10:00— 11:00h and again at 14:00-15:00h, lasting for about one hour. When approached, South Javan C. cursor readily flee under debris and /oomoea pes-caprae—Spinifex littoreus vegetation, as well as into the burrows of terrestrial crustaceans in sand and among plant roots. We noted that when sheltering inside burrows, C. cursor always turn their heads towards the burrow openings. Generally, C. cursor does not seek shelter above debris, but an observation in Batukaras showed that when there is nowhere else to hide, C. cursor may climb the base of a tree if it has crevices. Additionally, both ex-situ and in-situ observations showed that when cornered, C. cursor will bury themselves under the surface of loose sand for a short period of time or hide inside sand-filled coconut husks for longer periods. In captivity collected skinks exhibited sim- ilar behaviour during the day and returned to the surface later. In comparison, C. balinensis usually hides under leaf litter, branches, and bark while in captivity (Iskandar unpublished), but they never completely bury themselves in loose sand (Figure 3D). Lastly, we observed our captured specimens resting beneath sand during the night, but we could not confirm this behaviour in the wild. Future research investigating the genetic diversity of C. cursor from Java and beyond is recommended to determine whether these subpopu- lations represent separate species. ACKNOWLEDGEMENTS We thank Museum Zoologi Institut Teknologi Bandung for providing insights on surveying Cryptoblepha- rus along the southern coast of Java. We are also grateful for Balai Konservasi Sumber Daya Alam of the Special Region of Yogyakarta for issuing the permit Surat Angkut Tumbuhan dan Satwa - Dalam Negeri Check List 20 (4) - https://doi.org/10.15560/20.4.833 Sugianto et al. - New records of Cryptoblepharus cursor in Java 839 S.08/K.22-SKW./SATS-DN/1/2023 for the transfer specimens from Yogyakarta, Daerah Istimewa Yogyakarta to Bandung, Jawa Barat, Indonesia. ADDITIONAL INFORMATION Conflict of interest The authors declare that no competing interests exist. Ethical statement No ethical statement is reported. Funding This study was financially supported by the authors. Author contributions Conceptualization: DTI. Data curation: RAS, DTI. Formal analysis: RAS, DTI. Funding acquisition: RAS, DTI. Investigation: RAS, DTI, TSS. Methodology: RAS, DTI. Resources: RAS, DTI. Supervision: TSS. Visualization: RAS. Project administration: RAS, TSS. Software: RAS. Validation: DTI, TSS. Writing — original draft: RAS, DTI. Writing — review and editing: RAS, TSS. Author ORCID iDs Rizal Ananda Sugianto © https://orcid.org/0000-0001-6833-5809 Djoko Tjahjono Iskandar © https://orcid.org/0000-0002-8923-6081 Tati Suryati Syamsudin © https://orcid.org/0000-0003-2376-1593 Data availability All data that support the findings of this study are available in the main text. REFERENCES Akbar I (2017) iNaturalist observation of Lombok Snake-eyed Skink (Cryptoblepharus cursor). https://www.inaturalist.org/ observations/5980655. Accessed on: 2023-09-08. Asti H (2017) iNaturalist observation of Lombok Snake-eyed Skink (Cryptoblepharus cursor). https://www.inaturalist.org/ observations/5978111. Accessed on: 2024-04-04. Bagaskara SW (2017) iNaturalist observation of Lombok Snake-eyed Skink (Cryptoblepharus cursor). https://www. inaturalist.org/observations/919205/7. Accessed on: 2023-09-08. 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Previous records of Cryptoblepharus cursor in Indonesia No. Province Locality Geographic coordinates _Habitat Reference 1 Nusa Tenggara Barat Ampenan, Lombok Not provided Not described nor visible Barbour 1911 2 Sulawesi Selatan Pulau Samalona 05°08’S, 119°21’E Sandy beach with coral debris Horner 2007 4 Bali Unspecified Not provided Not described nor visible Iskandar unpublished 5 Jawa Timur Alas Purwo National Park Not provided Rocky beach Iskandar unpublished 6 Jawa Timur Pancur, Alas Purwo National Park 08°40'34"S, 114°22'21”E Rocks and driftwood Bagaskara 2017 7 08°40'37"S, 114°22’21"E Uncertain, not pictured Idrissa 2019 8 Plengkung, Alas Purwo National Park 08°43’58"S, 114°20’55”E Rocky beach Bajger 2022 9 Ngagelan, Alas Purwo National Park 08°37'18"S, 114°18’51”E Beach with herbaceous vegetation Supartha 2023a 10 Cemara, Banyuwangi 08°15'51"S, 114°22’20"E Coarse sand, various debris Supartha 2023b 11. Jawa Tengah Muara Puncu 07°51'17"S, 109°54’48"E Fine sands, wood debris, Spinifex littoreus Fathoni 2022 1 07°51'13"S, 109°54’41”E Fine sands, bamboo debris Dhiya’ulhag 2021a i) 07°51'13"S, 109°54’42”E Fine sands, bamboo debris Dhiya’ulhag 2021b 14 07°51'11"S, 109°54’41”E Fine sands, bamboo debris Dhiya’ulhag 2021c qe Roro Inten OF 5137S) 102" 50°33"E Fine sand Nurshafwan 2022 16 Daerah Istimewa Pasir Mendit Not provided Sand dunes Brongersma 1940 ig. oa 07°53'36"S, 110°0111"E Fine sand, Spinifex littoreus Fxpranoto 2017 18 07°53’36"S, 110°01'12”E Fine sand, Spinifex littoreus Akbar 2017 19 07°53’36"S, 110°01'11”E Fine sand, Spinifex littoreus Ramadani 2017 20 Maluku Pulau Leti 08°11'32”S, 127°41'10"E Uncertain, not shown Ramdani 2023 21 ‘Private location Not available Fine sand, Spinifex littoreus Asti 2017 Table A2. List of Cryptoblepharus cursor specimens examined in this study. Fieldtag _Locality Fieldtag _Locality RX-01 Congot, Daerah Istimewa Yogyakarta RY-02 Kadilangu, Daerah Istimewa Yogyakarta RX-02 Congot, Daerah Istimewa Yogyakarta RY-03 Kadilangu, Daerah Istimewa Yogyakarta RX-03 Congot, Daerah Istimewa Yogyakarta RY-04 Kadilangu, Daerah Istimewa Yogyakarta RX-04 Congot, Daerah Istimewa Yogyakarta RY-05 Kadilangu, Daerah Istimewa Yogyakarta RX-05 Congot, Daerah Istimewa Yogyakarta RY-06 Kadilangu, Daerah Istimewa Yogyakarta RX-06 Congot, Daerah Istimewa Yogyakarta RY-07 Kadilangu, Daerah Istimewa Yogyakarta RX-07 Congot, Daerah Istimewa Yogyakarta RY-08 Kadilangu, Daerah Istimewa Yogyakarta RX-08 Congot, Daerah Istimewa Yogyakarta RK-01 Batukaras, Jawa Barat RY-01 Kadilangu, Daerah Istimewa Yogyakarta Check List 20 (4) - https://doi.org/10.15560/20.4.833